Coming Soon:


Pfennig, D. W. & Martin, R. A. Evolution on an ecological time scale: a maternal effect mediates rapid population divergence and character displacement in spadefoot toads. (in review).


Martin, R. A. & Pfennig, D. W. Maternal induction of trophic polyphenism: implications for the evolution of novel resource-use phenotypes. (in review).


Pfennig, K. S. & Pfennig, D. W. Character displacement: Ecological and reproductive solutions to a common evolutionary problem. (in review).


Pfennig, D. W. & Kingsolver, J. G. 2008. Phenotypic selection. In S. A. Levin (ed.) Guide to Ecology, Princeton University Press, Princeton, NJ. (in press).


Published Articles:


Rice. A. M. & Pfennig, D. W. 2008. Analysis of range expansion in two species undergoing character displacement: Why might invaders generally “win” during character displacement? Journal of Evolutionary Biology 21: 696-704.


Ledon-Rettig, C., Pfennig, D. W., & Nascone-Yoder, N. 2008. Ancestral variation and the potential for genetic accommodation in larval amphibians: Implications for the evolution of novel feeding strategies. Evolution and Development 10: 316-325.


Harper, G. R. Jr. & Pfennig, D. W. 2008. Selection overrides gene flow to break down maladaptive mimicry. Nature 451: 1103-1106.


Pfennig, D. W. & Rice. A. M. 2007. An experimental test of character displacement’s role in promoting postmating isolation between conspecific populations in contrasting competitive environments. Evolution 61: 2433-2443. PDF


Harper, G. R. Jr. & Pfennig, D. W. 2007. Mimicry on the edge: Why do mimics vary in resemblance to their model in different parts of their geographical range? Proceedings of the Royal Society of London, Series B 274: 1955-1961. PDF


Kingsolver, J. G. & Pfennig, D.W. 2007. Patterns and power of phenotypic selection in nature. BioScience 57: 561-572. PDF


Pfennig, D. W., Rice. A. M., & Martin, R. A. 2007. Field and experimental evidence for competition’s role in phenotypic divergence. Evolution 61: 257-271. PDF


Rice. A. M. & Pfennig, D. W. 2007. Character displacement: in situ evolution of novel phenotypes or sorting of pre-existing variation? Journal of Evolutionary Biology 20: 448-459. PDF


Pfennig, D. W., Harper, G. R. Jr., Brumo, A. F., Harcombe, W. R., & Pfennig, K. S. 2007. Population differences in predation on Batesian mimics in allopatry with their model: selection against mimics is strongest when they are common. Behavioral Ecology and Sociobiology 61: 505-511. PDF


Pfennig, D. W., Rice. A. M., & Martin, R. A. 2006. Ecological opportunity and phenotypic plasticity interact to promote character displacement and species coexistence. Ecology 87: 769-779. PDF


Pfennig, K. S. & Pfennig, D. W. 2005. Character displacement as the ‘best of a bad situation’: fitness trade-offs resulting from selection to minimize resource and mate competition. Evolution 59: 2200-2208. PDF


Kingsolver, J. G. & Pfennig, D.W. 2004. Individual-level selection as a cause of Cope’s rule of phyletic size increase. Evolution 58: 1608-1612. PDF


Bull, J. J., Pfennig, D. W., & Wang, I-N. 2004. Genetic details, optimization and phage life histories. Trends in Ecology and Evolution 19: 76-82.


Pfennig, D. W. 2004. Putting genes in perspective. The American Scientist 92: 84-86. (review of Developmental Plasticity and Evolution, by M. J. West-Eberhard). PDF


Pfennig, D. W. & Murphy, P. J. 2003. A test of alternative hypotheses for character divergence between coexisting species. Ecology 84: 1288-1297. PDF


Reeve, H. K., Pfennig, D. W. (equal-authored) 2003. Genetic biases for showy males: are some genetic systems especially conducive to sexual selection? Proceedings of the National Academy of Sciences, USA 100: 1089-1094. PDF


Pfennig, D. W. & Murphy, P. J. 2002. How fluctuating competition and phenotypic plasticity mediate species divergence. Evolution 56: 1217-1228. PDF


Kingsolver, J. G., Pfennig, D. W., & Servedio, M. R. 2002. Migration, local adaptation, and the evolution of plasticity. Trends in Ecology and Evolution 17: 540-541.


Pfennig, D. W. 2002. Kin recognition. pp. 592-595. In M. Pagel (ed) Encyclopedia of Evolution, Oxford University Press, Oxford.


Frankino, W. A. & Pfennig, D. W. 2001. Condition-dependent expression of trophic polyphenism: effects of individual size and competitive ability. Evolutionary Ecology Research 3: 939-951. PDF


Pfennig, D. W., Harcombe, W. R., & Pfennig, K. S. 2001. Frequency-dependent Batesian mimicry. Nature 410: 323. PDF


Kingsolver, J. G. & Pfennig, D. W. 2001. Testing Darwin’s postulates. Trends in Ecology and Evolution 16: 165. (review of Adaptive Genetic Variation in the Wild, Edited by T. Mousseau, B. Sinervo, & J. A. Endler).


Pfennig, D. W. & Murphy, P. J. 2000. Character displacement in polyphenic tadpoles. Evolution 54: 1738-1749. PDF


Pfennig, D. W. 2000. Effect of predator-prey phylogenetic distance on the fitness consequences of predation: a tradeoff between nutrition and disease? The American Naturalist 155: 335-345. PDF


Loeb, M. L. G., Diene, L. & Pfennig, D. W. 2000. Egg dumping lace bugs preferentially oviposit with kin. Animal Behaviour 59: 379-383.


Pfennig, D. W. 2000. Attack of the tadpoles! Dragonfly 4: 16-17.


Pfennig, D. W., Collins, J. P., & Ziemba, R.E. 1999. A test of alternative hypotheses for kin recognition in cannibalistic tiger salamanders. Behavioral Ecology 10: 436-443. PDF


Pfennig, D. W. 1999. Cannibalistic tadpoles that pose the greatest threat to kin are most likely to discriminate kin. Proceedings of the Royal Society of London, Series B 266: 57-61. PDF


Hoffman, E. A. & Pfennig, D. W. 1999. Proximate causes of cannibalistic polyphenism in larval tiger salamanders. Ecology 80: 1076-1080.


Pfennig, D. W., Ho, S., & Hoffman, E. A. 1998. Pathogen transmission as a selective force against cannibalism. Animal Behaviour 55: 1255-1261.


Pfennig, D. W. 1998. The evolution of selflessness and selfishness. BioScience 48:9-11. (review of Survival Strategies: Cooperation and Conflict in Animal Societies, by R. Gadagkar).


Pfennig, D. W. & Frankino, W. A. 1997. Kin-mediated morphogenesis in facultatively cannibalistic tadpoles. Evolution 51: 1993-1999.


Pfennig, D. W. 1997. Kinship and cannibalism. BioScience 47: 667-675. PDF


Sherman, P. W., Reeve, H. K., & Pfennig, D. W. 1997. Recognition systems. pp. 69-96. In J. R. Krebs and N. B. Davies (eds) Behavioural Ecology: An Evolutionary Approach, 4th edn, Blackwell Scientific Publications, Oxford.


Pfennig, D. W. 1995. Absence of joint nesting advantage in desert seed harvester ants: evidence from a field experiment. Animal Behaviour 49: 567-575.



Pfennig, D. W. & Sherman, P. W. 1995. Kin recognition. Scientific American 272 (6): 98-103. PDF




Pfennig, D. W., Sherman, P. W., & Collins, J. P. 1994. Kin recognition and cannibalism in polyphenic salamanders. Behavioral Ecology 5: 225-232.


Pfennig, D. W. 1994. Cannibalism. Journal of Evolutionary Biology 4:121-123. (review of Cannibalism. Ecology and Evolution among Diverse Taxa, Edited by M. A. Elgar & B. J. Crespi).




Pfennig, D. W. & Collins, J. P. 1993. Kinship affects morphogenesis in cannibalistic salamanders. Nature 362: 836-838. PDF



Pfennig, D. W. & Reeve, H. K. 1993. Nepotism in a solitary wasp as revealed by DNA fingerprinting. Evolution 47: 700-704.


Pfennig, D. W., Reeve, H. K., & Sherman, P. W. 1993. Kin recognition and cannibalism in spadefoot toad tadpoles. Animal Behaviour 46: 87-94.


Pfennig, D. W. 1992. Polyphenism in spadefoot toad tadpoles as a locally-adjusted evolutionarily stable strategy. Evolution 46: 1408-1420. PDF


Pfennig, D. W. 1992. Proximate and functional causes of polyphenism in an anuran tadpole. Functional Ecology 6: 167-174. PDF


Pfennig, D. W. 1992. Social evolution of wasps. Journal of Evolutionary Biology 5: 729-731. (review of The Social Biology of Wasps, Edited by K. G. Ross & R. W. Matthews).


Pfennig, D. W. & Sherman, P. W. 1992. Identifying relatives. Science 255: 217-218. (review of Kin Recognition, Edited by P. G. Hepper).


Pfennig, D. W., Mabry, A., & Orange, D. 1991. Environmental causes of correlations between age and size at metamorphosis in Scaphiopus multiplicatus. Ecology 72: 2240-2248.


Pfennig, D. W., Loeb, M. L. G., & Collins, J. P. 1991. Pathogens as a factor limiting the spread of cannibalism among tiger salamanders. Oecologia 88: 161-166. PDF


Pfennig, D. W. 1990. “Kin recognition” among spadefoot toad tadpoles: a side-effect of habitat selection? Evolution 44: 785-798. PDF


Pfennig, D. W. 1990. Nestmate and nest discrimination among workers from neighboring colonies in social wasps, Polistes exclamans. Canadian Journal of Zoology 68: 268-271.


Pfennig, D. W. 1990. The adaptive significance of an environmentally-cued developmental switch in an anuran tadpole. Oecologia 85: 101-107.


Pfennig, D. W. & Reeve, H. K. 1989. Neighbor recognition and context-dependent aggression in a solitary wasp, Sphecius speciosus (Hymenoptera: Sphecidae). Ethology 80: 1-18.


Gamboa, G. J., Reeve, H. K., & Pfennig, D.W. 1986. The evolution and ontogeny of nestmate recognition in social wasps. Annual Review of Entomology 31: 431-454. PDF


Pfennig, D. W. & Klahn, J. E. 1985. Dominance as a predictor of cofoundress disappearance order in social wasps (Polistes fuscatus). Zeitschrift für Tierpsychologie (= Ethology) 67: 198-203.


Reeve, H. K., Shellman Reeve, J., & Pfennig, D. W. 1985. Eusociality and genetic variability: a re-evaluation. Evolution 39: 200-201.


Cothran, E. G., MacCluer, J. W., Weitkamp, L. R., Pfennig, D. W., & Boyce, A. J. 1984. Inbreeding and reproductive performance in Standardbred horses. Journal of Heredity 75: 220-224.


MacCluer, J. W., Boyce, A. J., Dyke, B., Weitkamp, L. R., Pfennig, D. W., & Parsons, C. J. 1983. Inbreeding and pedigree structure in Standardbred horses. Journal of Heredity 74: 394-399.


Pfennig, D. W., Gamboa, G. J., Reeve, H K., Shellman Reeve, J., & Ferguson, I. D. 1983. The mechanism of nestmate discrimination in social wasps (Polistes, Hymenoptera: Vespidae). Behavioral Ecology and Sociobiology 13: 299-305.


Pfennig, D. W., Reeve, H. K., & Shellman, J. S. 1983. Learned component of nestmate discrimination in workers of a social wasp, Polistes fuscatus (Hymenoptera: Vespidae). Animal Behaviour 31: 412-416.